last updated: 15.IX.2009



Pipunculidae literature and identification keys focusing on the European fauna

During the last 25 years, the knowledge about Pipunculidae has increased significantly. Many genera from different zoogeographical regions have been revised, resulting in the description of many new species.
Here, you can find a compilation of mostly taxonomic literature, necessary for identification purposes. The focus of this listing has been put on the European fauna.



Identification key to European genera/tribes of Pipunculidae


Source: Dunk (1997); ammended.



Faunistics


Phylogeny

  • De Meyer, M. (1993): Phylogeny and evolutionary zoogeography of the Hawaiian Pipunculidae (Diptera).- Z. zool. Syst. Evolut.-forsch. 31: 119-126.
  • De Meyer, M. (1994): Phylogenetic relationships within the Cephalopsini (Diptera, Pipunculidae).- Bull. Annls Soc. r. belge Ent. 130: 7-18.
  • De Meyer, M. (1996): Cladistic and biogegraphic analyses of Hawaiian Pipunculidae (Diptera) revisited.- Cladistics 12: 291-303.
  • Kehlmaier, C. & Assmann, T. (2010): Molecular analysis meets morphology-based systematics — a synthetic approach for Chalarinae (Insecta: Diptera: Pipunculidae).- Systematic Entomology 35: 181-195. DOI: 10.1111/j.1365-3113.2009.00500.x
  • Rafael, J.A. (1988): [Considerations about the phylogeny of Nephrocerinae and description of the male of Protonephrocerus chiloensis Collin, 1931 (Diptera, Pipunculidae)].- Revta bras. Ent. 32 (3/4): 465-470.
    Note: The article is in Portuguese, including an English summary.
  • Rafael, J.A. & De Meyer, M. (1992): Generic classification of the family Pipunculidae (Diptera): a cladistic analysis.- Journal of Natural History 26: 637-658.
  • Skevington, J.H. (2006): Revision of Fijian Collinias Aczél (Diptera: Pipunculidae).- Fiji Arthropods V. Bishop Museum Occasional Papers 89: 13-43.
  • Skevington, J.H. & Yeates, D.K. (2000): Phylogeny of the Syrphoidea (Diptera) inferred from mtDNA Sequences and Morphology with Particular Reference to Classification of the Pipunculidae (Diptera).- Molecular Phylogenetics and Evolution 16 (2): 212-224.
  • Skevington, J.H. & Yeates, D.K. (2001): Phylogenetic classification of Eudorylini (Diptera: Pipunculidae).- Systematic Entomology 26: 421-452.
    Note: This work includes the description of two new genera and the latest key to the world genera of extant Pipunculidae.
  • Skevington, J.H.; Kehlmaier, C. & Stĺhls, G. (2007): Molecular barcoding: Mixed results for Pipunculidae (Diptera).- Zootaxa 1423: 1-26; Brisbane.
  • Stĺhls, G.; Hippa, H.; Rotheray, G.; Muona, J. & Gilbert, F. (2003): Phylogeny of Syrphidae (Diptera) inferred from combined analysis of molecular and morphological characters.- Systematic Entomology 28: 433-450.
    Note: One species of Pipunculidae (Jassidophaga villosa) is used as outgroup.


The Genera of Pipunculidae


Subfamily Chalarinae

Chalarus Walker, 1834

    Europe
  • Jervis, M.A. (1992): A taxonomic revision of the pipunculid fly genus Chalarus Walker, with particular reference to the European fauna.- Zoological Journal of the Linnean Society 105: 243-352; London.
  • Kehlmaier, C. & Assmann, T. (2008): The European species of Chalarus WALKER, 1834 revisited (Diptera: Pipunculidae).- Zootaxa 1936: 1-39; Brisbane.
Additional literature:
  • Coe, R.L. (1966b): Some British species of Chalarus and Verrallia (Diptera: Pipunculidae).- Proc. R. ent. Soc. Lond. (B) 35 (11-12): 149-160.
  • Dempewolf, M. (1998): On the taxonomy of two German pipunculid flies with remarks on the terminology of the male genitalia.- Beiträge zur Entomologie 48 (2): 531-537; Berlin.
  • Kehlmaier, C. (2003d): Pipunculidae (Diptera) from a forest ecosystem in northern Spain, with the description of a new species.- Entomologische Zeitschrift 113 (3): 87-94; Stuttgart.

  • Jassidophaga Aczél, 1939

      Europe
    • Kehlmaier, C. (2006): The West-Palaearctic species of Jassidophaga Aczél and Verrallia Mik described up to 1966 (Diptera: Pipunculidae).- Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 697, 34pp.; Stuttgart.
    • Kuznetzov, S. Y. (1992): The Palaearctic species of the genus Verrallia (Diptera: Pipunculidae).- Zoosystematica Rossica 1: 102-116; St. Petersburg.
    Additional literature prior to the above mentioned:
  • Coe, R.L. (1966a): Pipunculidae.– Handbook for the Identification of British Insects, Vol. X (Part 2c): 1-83; London.
  • Coe, R.L. (1966b): Some British species of Chalarus and Verrallia (Diptera: Pipunculidae).- Proc. R. ent. Soc. Lond. (B) 35 (11-12): 149-160.
  • Lauterer, P. (1981): Contribution to the knowledge of the family Pipunculidae of Czechoslovakia (Diptera).– Acta Musei Moraviae, Scientiae naturales 66: 123-150; Brno

  • Verrallia Mik, 1899

      Europe
    • Kehlmaier, C. (2006): The West-Palaearctic species of Jassidophaga Aczél and Verrallia Mik described up to 1966 (Diptera: Pipunculidae).- Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 697, 34pp.; Stuttgart.
    • Kuznetzov, S. Y. (1992): The Palaearctic species of the genus Verrallia (Diptera: Pipunculidae).- Zoosystematica Rossica 1: 102-116; St. Petersburg.



    Subfamily Nephrocerinae

    Nephrocerus Zetterstedt, 1838

      Europe
    • Grootaert, P. & De Meyer, M. (1986): On the taxonomy and ecology of Nephrocerus Zetterstedt (Diptera, Pipunculidae) with a redescription of N. lapponicus and a key to the European species.- Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, Entomologie 56: 85-91.


    Protonephrocerus Collin, 1931
    Not present in Europe.



    Subfamily Pipunculinae

    Tribe Pipunculini

    Parapipunculus Rafael, 1986

    Not present in Europe and considered synonymus with Pipunculus by Skevington, J. & Marshall, S.A. (1998: 2).


    Pipunculus Latreille, 1802

      Europe
    • Kehlmaier, C. (2008): Finnish Pipunculidae (Diptera) Studies Part I: Taxonomic Notes on Cephalops Fallén, 1810, Pipunculus Latreille, 1802 and Tomosvaryella Aczél, 1939.- Zootaxa 1672: 1-42; Brisbane.
    • Kuznetzov, S. Y. (1991): New Palaearctic species of the genus Pipunculus (Diptera, Pipunculidae).- Dipterological Research 2: 1-52.
    • Skevington, J. & Marshall, S. A. (1998): Systematics of New World Pipunculus (Diptera: Pipunculidae).- Thomas Say Publications in Entomology: Monographs, 201pp.; Lanham.
    Additional literature prior to the above mentioned work:
  • Földvári, M. & Kozánek, M. (2001): New morphological and faunistic records of Hungarian Pipunculidae (Diptera).- Folia entomologica hungarica Rovartani Közlemények 62: 293-303.
  • Kozánek, M. (1981a): Genus Pipunculus Latreille (Diptera, Pipunculidae) in Czechoslovakia.- Annotationes Zoologicae et Botanicae 142: 1-16; Bratislava.
  • Kozánek, M. (1981b): Description of a new species of Pipunculus Latr. (Diptera, Pipunculidae) and a redescription of P. omissinervis Beck.- Biológia 36 (8): 605-609; Bratislava.
  • Kuznetzov, S. Y. (1993): A new Pipunculus species from Spain (Diptera: Pipunculidae).- Dipterological Research 4 (3): 139-144.


  • Tribe Cephalopsini

    Cephalops Fallén, 1810

      Europe
    • Ackland, D. M. (1993): Notes on British Cephalops Fallén, 1810 with description of a new species, and Microcephalops De Meyer, 1989, a genus new to Britain (Dipt., Pipunculidae).- Entomologist's Monthly Magazine 129: 95-105.
    • De Meyer, M. (1989a): The West-Palaearctic species of the pipunculid genera Cephalops and Beckerias (Diptera): classification, phylogeny and geographical distribution.- Journal of Natural History 23: 725-765.
    • De Meyer, M. (1989b): Systematics of the Nearctic species of the genus Cephalops Fallén (Diptera, Pipunculidae).- Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, Entomologie 59: 99-130; Brussel.
      Note: Redescription of C. varius (Cresson, 1911), only recently recorded from Europe.
    • Kehlmaier, C. & De Meyer, M. (2005): On the identity of Pipunculus straminipes BECKER, 1900 (Diptea: Pipunculidae).- Studia dipterologica 11(2) [2004]: 600-602; Halle an der Saale.


    Cephalosphaera Enderlein, 1936

      Europe
    • De Meyer, M. (1989): The West-Palaearctic species of the pipunculid genera Cephalops and Beckerias (Diptera): classification phylogeny and geographical distribution.- Journal of Natural History 23: 725-765.



    Tribe Microcephalopsini

    Collinias Aczél, 1940

    Not present in Europe.


    Microcephalops De Meyer, 1989

      Europe
    • De Meyer, M. (1989a): The West-Palaearctic species of the pipunculid genera Cephalops and Beckerias (Diptera): classification phylogeny and geographical distribution.- Journal of Natural History 23: 725-765.
    • Dunk, K. von der & Lauterer, P. (1998): More details on the rare Eudorylas furvulus Collin and Microcephalops opacus (Fallén) comb. n. (Diptera: Pipunculidae).- Acta Mus. Moraviae, Sci. biol. 82: 163-171.
      Note: The article gives a redescription of the only European species of this genus.



    Tribe Eudorylini

    Allomethus Hardy, 1943

    Not present in Europe.


    Amazunculus Rafael, 1986

    Not present in Europe.


    Basileunculus Rafael, 1987

    Not present in Europe.


    Claraeola Aczél, 1940

      Europe
    • Kehlmaier, C. (2005a): Taxonomic revision of European Eudorylini (Insecta, Diptera, Pipunculidae).- Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 41: 45-353; Hamburg.


    Clistoabdominalis Skevington, 2001

      Europe
    • Kehlmaier, C. (2005a): Taxonomic revision of European Eudorylini (Insecta, Diptera, Pipunculidae).- Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 41: 45-353; Hamburg.


    Dasydorylas Skevington, 2001

      Europe
    • Kehlmaier, C. (2005a): Taxonomic revision of European Eudorylini (Insecta, Diptera, Pipunculidae).- Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 41: 45-353; Hamburg.


    Elmohardyia Rafael, 1987

    Not present in Europe.


    Eudorylas Aczél, 1940

      Europe
    • Kehlmaier, C. (2005a): Taxonomic revision of European Eudorylini (Insecta, Diptera, Pipunculidae).- Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 41: 45-353; Hamburg.
    Additional literature prior to the above mentioned revision:
  • Ackland, D. M. (1994): Draft key to Eudorylas.- Unpublished manuscript.
  • Ackland, D. M. (1999): A new species of Eudorylas from Scotland (Diptera, Pipunculidae).- Dipterist's Digest 6: 35-39.
  • Ackland, D.M. (2002): Test keys to British Eudorylini.– The Piercer. A Pipunculidae Newsletter, 3, Dipterists Forum, Pipunculidae Study Group (Internet: http://www.dipteristsforum.org.uk).
  • Becker, T. (1897): Dipterologische Studien V. Pipunculidae.- Berliner Entomologische Zeitschrift 42 (1/2): 25-100.
    Note: The article is in German.
  • Becker, T. (1900): Dipterologische Studien V. Pipunculidae. Erste Fortsetzung.- Berliner Entomologische Zeitschrift 45: 215-252.
    Note: The article is in German.
  • Becker, T. (1921): Neue Dipteren meiner Sammlung. Pipunculidae (Dorylidae).- Wiener Entomologische Zeitung 38: 123-132 & 149-167.
    Note: The article is in German.
  • Coe, R. L. (1966): Pipunculidae.- In: Handbook for the Identification of British Insects Vol. X, Part 2 (c): 83pp.
  • De Meyer, M. (1993): Notes on the Pipunculidae fauna (Diptera) of Switzerland.- Mitteilungen der schweizerischen entomologischen Gesellschaft 66: 41-52.
    Note: Description of Eudorylas kozaneki.
  • De Meyer, M. (1995): The pipunculid flies of Israel and the Sinai (Insecta, Diptera, Pipunculidae).- Spixiana 18 (3): 283-319; Munich.
  • De Meyer, M. (1997): Contribution to the Pipunculidae fauna of Spain (Diptera).- Beiträge zur Entomologie 47 (2): 421-450; Berlin.
  • Dempewolf, M. (1996): Taxonomical notes on west palaearctic Pipunculidae including the description of two new Eudorylas species (Diptera).- Studia dipterologica 3 (1): 87-92; Halle an der Saale.
  • Dempewolf, M. (1998): On the taxonomy of two German pipunculid flies (Diptera) with remarks on the terminology of the male terminalia.- Beiträge zur Entomologie 48 (2): 531-537.
  • Dempewolf, M. & Dunk, K. von der (1996): A new Eudorylas species from Southern and Western Germany (Diptera, Pipunculidae).- Studia dipterologica 3 (1): 93-96; Halle an der Saale.
  • Duda, O. (1940): [New and insufficiently known Palaearctic Diptera from my own collection].- Folia Zoologica et Hydrobiologica 10 (1): 214-226.
    Note: Description of E. dudai (Hardy, 1947) (as Dorylas curvitibia) and E. electus (Hardy, 1947) (as Dorylas caudatus). The article is in German.
  • Dunk, K. von der (1995): Description of two new Eudorylas species and the so far unknown female of E. restrictus Coe, 1966 (Dipt.- Pipunculidae).- galathea 11 (3): 119-125; Nürnberg.
  • Dunk, K. von der (1997): Key to Central European species of Pipunculidae (Diptera).- Studia dipterologica 4 (2): 289- 335; Halle an der Saale.
  • Dunk, K. von der & Lauterer, P. (1998): More details on the rare Eudorylas furvulus Collin and Microcephalops opacus (Fallén) comb. n. (Diptera: Pipunculidae).- Acta Mus. Moraviae, Sci. biol. 82: 163-171.
  • Janssens, E. (1955): Mission E. Janssens & R. Tollet en Crece (Juillet-Aout 1953) 13e Note. Diptera Pipunculidae.- Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 31: 1-6.
    Note: Description of E. straeleni. The article is in French.
  • Kozánek, M. (1988): Description of two new species of the genus Eudorylas (Diptera, Pipunculidae) from Soviet Middle Asia.- Biológia 43 (10): 945-948; Bratislava.
  • Kozánek, M. (1993): Description of five new Eudorylas Aczél species from Central Europe (Diptera: Pipunculidae).- Entomological Problems 24 (1): 39-50; Bratislava.
  • Kozánek, M. & Kehlmaier, C. (2004): Pipunculidae of Slovakia: additions and corrections to faunal list, with a description of a new Eudorylas (Diptera).- Entomological Problems 34 (1-2): 21-36; Bratislava.
  • Kuznetzov, S. Y. (1990): New Palaearctic species of Pipunculidae (Diptera).- Dipterological Research 1: 23-50.
    Note: Description of E. rusticus.
  • Kuznetzov, S. Y. (1995): Eudorylas electus (Hardy): new record and redescription (Diptera, Pipunculidae).- Acta Zool. Fennica 199: 33-35; Helsinki.


  • Tribe Tomosvaryellini

    Dorylomorpha Aczél, 1939

      Europe
    • Albrecht, A. (1990): Revision, phylogeny and classification of the genus Dorylomorpha (Diptera, Pipunculidae).- Acta Zoologica Fennica 188: 1-240; Helsinki.
    • Kehlmaier, C. (2008b): 4.3.19 Pipunculidae.- In: Ziegler, J. (ed.): Diptera Stelviana. A dipterological perspective on a changing alpine landscape. Volume 1.- Studia dipterologica Supplement 16: 207-220 pp.; Ampyx-Verlag: Halle (Saale).


    Tomosvaryella Aczél, 1939

      Europe
    • Coe, R.L. (1969): Some Pipunculidae (Diptera) from Southern Spain, with description of a new species.- Entomologiske Meddelelser 37: 3-8.
    • De Meyer, M. (1995): The pipunculid flies of Israel and the Sinai (Insecta, Diptera, Pipunculidae).- Spixiana 18 (3): 283-319; Munich.
    • De Meyer, M. (1997): Contribution to the Pipunculidae fauna of Spain (Diptera).- Beiträge zur Entomologie 47 (2): 421-450; Berlin.
    • De Meyer, M.; Földvári, M. & Báez, M. (2000): The Pipunculidae (Diptera) fauna of the Canary Islands and Madeira.- Bulletin S.R.B.E./K.B.V.E. 136: 144-152.
      Note: Taxonomic comments on T. ornatipes stat. rev. and T. glabrum stat. rev. (with T. tecta syn. nov.).
    • Földvári, M. & De Meyer, M. (1999): Revision of central and west European Tomosvaryella Aczél species (Diptera, Pipunculidae).- Acta Zoologica Scientiarum Hungaricae 45 (4): 299-334; Budapest.
    • Janssens, E. (1955): Mission E. Janssens & R. Tollet en Crece (Juillet-Aout 1953) 13e Note. Diptera Pipunculidae.- Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 31: 1-6.
      Note: Description of T. olympicola.
    • Kehlmaier, C. (2008): Finnish Pipunculidae (Diptera) Studies Part I: Taxonomic Notes on Cephalops Fallén, 1810, Pipunculus Latreille, 1802 and Tomosvaryella Aczél, 1939.- Zootaxa 1672: 1-42; Brisbane.
    • Kuznetzov, S. Y. (1993): A new European species of Tomosvaryella (Diptera, Pipunculidae).- Zoosystematica Rossica 2: 177-178; St. Petersburg.
    • Kuznetzov, S. Y. (1994): New Palaearctic species of Tomosvaryella Aczél (Diptera, Pipunculidae) with redescription of T. rondanii (Collin).- Dipterological Research 5: 107-207.



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